Research Plant systematics / Systematics of Orchidaceae and Liliaceae / Population Genetics of Orchidaceae and Liliaceae, especially Goodyera, ● Liparis, Veratrum etc.
Research Molecular phylogeny of metazoan animal groups (Nematoda, Mollusca, Acanthocephala) / Evolutionary genomics of metazoan groups using NGS (Next-Generation Sequencing) / Taxonomy of nematodes and mollusks (Morphology)
